Born in 1922 as the youngest of five children in a rural village that is now part of eastern Pakistan, Khorana learned to read and write with help from his father, according to the Nobel Prize’s biography of the biochemist. With a number of scholarships, Khorana went on to earn a doctorate in organic chemistry in 1948. He conducted his Nobel Prize-winning research on nucleotides at the University of Wisconsin, and he later became the Alfred P. Sloan Professor of Biology and Chemistry at the Massachusetts Institute of Technology.
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